An Early Cretaceous non-avian dinosaur and bird footprint assemblage from the Laiyang Group in the Zhucheng basin, Shandong Province, China.

Diverse non-avian dinosaur (sauropod and ornithopod) and bird tracks occur in the Lower Cretaceous Yangjiazhuang Formation at the Zhangzhuhewan Village track site in Zhucheng basin, Shandong Province, China. Non-avian dinosaur tracks at this tracksite are heavily deformed, having been registered in waterlogged sediment. Sauropod pes and manus impressions, made by a slowly advancing track maker, are roughly equal in area. The ornithopod tracks are of the generalized Iguanodon-hadrosaur morphotype. The bird tracks are unlike other Mesozoic bird tracks from China, but are similar to the shorebird-like ichnotaxon Jindongornipes. Among tracks previously reported from the Laiyang Group, Laiyangpus is of uncertain affinity and its type specimen has been lost, and Paragrallator is referable to Anchisauripus isp. The new tracks described herein further increase the known dinosaur diversity in eastern Shandong Province.