中国发现世界上最原始的鸟类化石:华美金凤鸟(新属、新种)

    First avialian birdfrom China(Jinfengopteryx elegans gen. et sp. nov.)

    • 摘要: 始祖鸟的发现被认为是19世纪最重要的科学发现之一,迄今为止共发现了7枚保存骨骼、羽毛印痕的化石标本和1枚单根羽毛化石标本。由于始祖鸟既显示了明显的爬行动物的特征又保存了精美的羽毛,140多年来人们一直将其作为介于恐龙与鸟类之间的“中间环节”(“missing link”),但更倾向认为它是世界上最古老的鸟,或鸟类的始祖。由于始祖鸟化石稀少,加之地理分布十分局限,长期以来国际上一直围绕鸟类起源问题展开了激烈的争论,提出了各种各样的假说。直到20世纪90年代,在中国辽西晚中生代地层中发现了大量长羽毛的恐龙和原始鸟类化石,有力地支持了鸟类起源于小型兽脚类恐龙的学术观点,使赫 140多年前提出的“假说”成为当今国际科学界占主流地位的学说和理论,基本解决了长期困扰国际科学界的鸟类起源问题,遗憾的是,在过去的几年中,中国一直没有发现与德国始祖鸟十分接近或与德国始祖鸟处于同样进化水平的原始鸟类化石,这种缺憾使我们难以对鸟类定义问题进行深入的研究和探讨,2002年,笔者等在中国辽西早白垩世断层中发现了2种初鸟类化石,即中华神州鸟和东方吉祥鸟。通过比较解剖学研究和支序分析,这2种初鸟类要比始祖鸟略微进步一些,在研究鸟类的早期演化方面具有重要意义;但仍然难以对鸟类定义问题作进一步研究。本文首次报道了一件采自中国河北省丰宁县龙凤山桥头组的初鸟类化石——华美金凤鸟(新属、新种)(Jinfengopteryx elegans gen.et sp.nov),与其共生的化石有Lycoptera davidi,Peipiaosteus pani,yanosteus longidorsalis(鱼类);Jibeinia luanhera,Protopteryx fengningensis(鸟类);Ephemeropsis trisetalis(昆虫)及少量植物。支序分析表明,华美金凤鸟处于初鸟类谱系树的基部,比始祖鸟略微原始,与后者为姊妹群关系。奔龙等兽脚类恐龙是它们的外类群。金凤鸟、始祖鸟是中华神州鸟等原始鸟类的外类群。这说明华美金凤鸟处在恐龙向鸟类演化过程中一个非常关键的位置,是世界上迄今所发现的最原始的初鸟类。华美金凤鸟的发现在研究鸟类起源、恐龙-鸟类的系统关系等方面具有重要的科学意义。

       

      Abstract: Archaeopteryx has been recognized as the earliest known bird in the world for more than 140 years, and it is well documented from seven skeletons with feather impressions and an isolated feather. Archaeopteryx shows a combination of diapsid ("reptile-like") plesiomorphies and the derived ("bird-like") features. It provides the most important evidence for the hypothesis that birds originated from a dinosaurian ancestor. Since 1990s, the rich assemblage of taxonomically diverse and numerically abundant feathered theropods and primitive birds have been found from the Early Cretaceous Yixian Formation of western Liaoning, China. These findings further strengthen the hypothesis that birds were derived from small theropod dinosaurs. However, no avialian birds were found from China at that time. Until 2002, two avialian birds, Shenzhouraptor sinensis and Jixiangornis orientalis, were reported from the Early Cretaceous beds in western Liaoning, China. Here we describe a new well-preserved avialian bird from northern Hebei, China. Our phylogenetic analyses have placed it at the root of the avialian family tree. It sheds new light on the phylogenetic relationship between non-avian theropods and avian theropods (birds) and on the definition of birds.

       

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